Scorecard
Watson shared a solid opening partnership of 84 with Chris Rogers to give the touring side a good start in their bid to score 311 runs for victory.
But he perished immediately after the drinks break for 46, trapped lbw by a full-length delivery from Stuart Broad.
Cowan fell to part-time spinner Joe Root just before the interval, leaving Rogers unbeaten on 50.
Earlier, Ian Bell made 109 and Broad 65 before England were bowled out for 375 in their second innings.
Bell and Broad, resuming on 326 for six, quickly reached the individual milestones their tenacious partnership deserved.
Mitchell Starc got Australia's day off to a bad start with a wild beamer which went through first slip to the boundary and Broad slashed James Pattinson for four to get to fifty.
Bell pushed Starc for a single to post his 18th Test century, a marathon effort of intense concentration lasting more than six hours.
Bell, often criticised for failing to deliver under pressure, leapt up and punched the air after completing his run before raising his bat to all sides of the ground.
The seventh-wicket partnership of 138 ended when Broad edged Pattinson through to wicketkeeper Brad Haddin.
The England fast bowler walked straight off to ironic cheers after the controversial incident on Friday when he edged Ashton Agar to slip and stayed at the wicket having being given not out by the umpire.
FINE DELIVERY
Bell drove Starc to the cover boundary and chopped his next ball for four but he nicked a fine delivery from Starc through to Haddin and left the field to a standing ovation.
He faced 267 balls, hit 15 fours and batted for six-and-a-half hours in an innings which put England firmly in control of the match.
Australia wrapped up the England tail quickly.
Graeme Swann, on nine, edged Peter Siddle to Michael Clarke at slip and James Anderson lasted only two balls before he was caught by Phil Hughes at mid-wicket off the persevering Siddle.
Watson and Rogers survived the seven overs before lunch without any major alarms to reach the interval on 28 for no wicket.
The pair also batted confidently through the first hour of the afternoon session as England struggled for inspiration under blue skies.
Watson peppered the boundary with eight fours and he looked certain to reach his fifty before a lapse in concentration gave Broad England's first breakthrough.
Watson referred the umpire's decision to give him out but replays showed the ball would have hit the stumps and he left the field shaking his head after once again failing to turn a good start into a really significant contribution for his team.
Cowan, on a king pair, got off the mark with a cut for four off Steven Finn before the 35-year-old Rogers reached his first Test fifty, off 104 balls and including eight fours.
Cowan was growing in confidence but, on 14 and just three balls before tea, he tried to drive a wide ball from Root and edged a simple catch to Jonathan Trott at slip.
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